Development of the modern, highly specialised and economically efficient, forestry in the 1700s has substantially changed the structural characteristics and dynamics of European woodlands. In particular, the ban on forest grazing that had been almost universally adopted throughout all Central Europe, unleashed the processes of development of shade−tolerant undergrowth, leading to the loss of semi−open woods and to functional, ecological, isolation of forests from the wider landscape context. In a separate development, due the recent socio−economic changes, scores of farmland have been abandoned, turning variegated countryside to feral landscapes, increasingly covered by woody vegetation. Although such changes are in line with recommendations focused on the climate issues, we believe that the ‘ecological intensification’ of the abandoned landscapes may provide more benefits than their permanent abandonment. In order to verify that intuition, we studied ecological features and silvopastoral benefits of a landscape unit in the Carpathian foothills of SE Poland, which, during the last twelve years has been grazed by cattle, let in after the former ten years since the agricultural abandonment. We provide the information on the landscape’s mosaic structure with regard to the habitats use by cattle. In addition, we investigated the animals’ reaction to the exposure of the local arboreal forage, of which the chemistry and nutritional values have been compared to those of the herbaceous one. As shown by the study, cattle can be used as a factor successfully preventing succession and sustaining the balance between wooded and non−wooded habitats. We revealed that the wood−pasture’s tree hay was very attractive to cattle, compensating deficits of the herbaceous fodder. Our study proved that the grazing herd of robust cattle breeds can turn a feral, post−agricultural land into a working wood−pasture, consisting of interconnected open grasslands and various facets of woodland, some of them missing in contemporary high forests. Such landscapes can provide multiple ecosystem services, scarce or absent in abandoned, overgrown farmlands. We advocate, therefore, for the transition management of abandoned farmlands towards the integrated silvopastoral land use, contributing to the land’s economic value and landscape’s ecosystem services, and the restoration of woodland’s characteristics, disappeared in contemporary forests.

Affek, A.N., Wolski, J., Zachwatowicz, M., Ostafin, K., Radeloff, V.C., 2021. Effects of post-WWII forced displacements on long-term landscape dynamics in the Polish Carpathians. Landscape and Urban Planning, 214: 104164. DOI: https://doi.org/10.1016/j.landurbplan.2021.104164.
AOAC, 2020. Official methods of analysis of association of official analytical chemists (AOAC). Available from: http://www.eoma.aoac.org/.
Appleton, J., 1975. The experience of landscape. London: Wiley and Sons, 293 pp.
Arcanum-maps, 2022. Europe in the XIX Century. Budapest: Arcanum Adatbázis Kft. Available from: https://tinyurl.com/arcanummaps.
Bakshi, M.P.S., Wadhwa, M., 2012. Comparative nutritional evaluation of forest tree leaves and wild grasses of semi hilly arid zone in Punjab. Indian Journal of Animal Sciences, 82 (8): 918. DOI: https://doi.org/10.56093/ijans.v82i8.23038.
Biró, M., Molnár, Z., Öllerer, K., Lengyel, A., Ulicsni, V., Szabados, K., Kiš, A., Perić, R., Demeter, L., Babai, D., 2020. Conservation and herding co-benefit from traditional extensive wetland grazing. Agriculture, Ecosystems and Environment, 300: 106983. DOI: https://doi.org/10.1016/j.agee.2020.106983.
Bobiec, A., Kuijper, D.P.J., Niklasson, M., Romankiewicz, A., Solecka, K., 2011. Oak (Quercus robur L.) regeneration in early successional woodlands grazed by wild ungulates in the absence of livestock. Forest Ecology and Management, 262 (5): 780-790. DOI: https://doi.org/10.1016/j.foreco.2011.05.012.
Bobiec, A., Paderewski, J., Gajdek, A., 2021. Urbanisation and globalised environmental discourse do not help to protect the bio-cultural legacy of rural landscapes. Landscape and Urban Planning, 208: 104038. DOI: https://doi.org/10.1016/j.landurbplan.2021.104038.
Bobiec, A., Podlaski, R., Ortyl, B., Korol, M., Havryliuk, S., Öllerer, K., Ziobro, J.M., Pilch, K., Dychkevych, V., Dudek, T., Mázsa, K., Varga, A., Angelstam, P., 2019. Top-down segregated policies undermine the maintenance of traditional wooded landscapes: evidence from oaks at the European Union’s eastern border. Landscape and Urban Planning, 189: 247-259. DOI: https://doi.org/10.1016/j.landurbplan.2019.04.026.
Bobiec, A., Reif, A., Öllerer, K., 2018. Seeing the oakscape beyond the forest: a landscape approach to the oak regeneration in Europe. Landscape Ecology, 33: 513-528. DOI: https://doi.org/10.1007/s10980-018-0619-y.
Brambilla, M., Gustin, M., Vitulano, S., Falco, R., Bergero, V., Negri, I., Bogliani, G., Celada, C., 2017. Sixty years of habitat decline: impact of land-cover changes in northern Italy on the decreasing ortolan bunting Emberiza hortulana. Regional Environmental Change, 17: 323-333. DOI: https://doi.org/10.1007/s10113-016-1019-y.
Braun-Blanquet, J., Pavillard, J., 1925. Vocabulaire de sociologie végétale. Montpellier: Roumégous et Déhan, 25 pp.
von Carlowitz, H.C., 1713. Sylvicultura Oeconomica, oder Haußwirthliche Nachricht und Naturmäßige Anweisung zur Wilden Baum Zucht. Leipzig: Joachim Hamberger, 640 pp. Available from: https://www.amazon.pl/s/ref=dp_byline_sr_book_1?ie=UTF8&field-author=Joachim+Hamberger&search-alias=books.
Dzwonko, Z., Gawroński, S., 1994. The role of woodland fragments, soil types, and dominant species in secondary succession on the western Carpathian foothills. Vegetatio, 111: 149-160. DOI: https://doi.org/10.1007/BF00040334.
EC (European Commission), 2021. A European Green Deal: Striving to be the first climate-neutral continent. Available from: https://ec.europa.eu/info/strategy/priorities-2019-2024/european-green-deal/delivering-european-green-deal_en.
Faliński, J.B., 1986. Vegetation dynamics in temperate lowland primeval forests, ecological studies in Białowieża Forest. Dordrecht: Dr W Junk Publishers, 537 pp.
González-Hernández, M.P., Karchesy, J., Starkey, E.E., 2003. Research observation: Hydrolyzable and condensed tannins in plants of northwest Spain forests. Journal of Range Management, 56: 461-465. DOI: https://doi.org/10.2307/4003837.
Gowda, N.K.S., Ramana, J.V., Prasad, C.S., Singh, K., 2004. Micronutrient content of certain tropical conventional and unconventional feed resources of southern India. Tropical Animal Health and Production, 36: 77-94. DOI: https://doi.org/10.1023/B:TROP.0000009522.30949.1d.
Gough, LA., Birkemoe, T., Sverdrup-Thygeson, A., 2014. Reactive forest management can also be proactive for wood-living beetles in hollow oak trees. Biological Conservation, 180: 75-83. DOI: https://doi.org/10.1016/j.biocon.2014.09.034.
Grainger, C., Clarke, T., Auldist, M.J., Beauchemin, K., McGinn, S., Waghorn, G.C., Eckard, R., 2009. Potential use of Acacia mearnsii condensed tannins to reduce methane emissions and nitrogen excretion from grazing dairy cows. Canadian Journal of Animal Science, 89: 241-251. DOI: https://doi.org/10.4141/CJAS08110.
Holzl, R., 2010. Historicizing sustainability: German scientific forestry in the eighteenth and nineteenth centuries. Science as Culture, 19 (4): 431-460. DOI: https://doi.org/10.1080/09505431.2010.519866.
Horak, J., Vodka, S., Kout, J., Halda, J.P., Bogusch, P., Pech, P., 2014. Biodiversity of most dead wood-dependent organisms in thermophilic temperate oak woodlands thrives on diversity of open landscape structures. Forest Ecology and Management, 315: 80-85. DOI: https://doi.org/10.1016/j.foreco.2013.12.018.
Jaroszewicz, B., Jankowska-Błaszczuk, M., Żmihorski, M., Hałatkiewicz, T., 2020. An efficient tool for the maintenance of thermophilous oak forest understory – Sheep or brush cutter? Forests, 11 (5): 582. DOI: https://doi.org/10.3390/f11050582.
Jaworski, A., Kołodziej, Z., Porada, K., 2002. Structure and dynamics of stands of primeval character in selected areas of the Bieszczady National Park. Journal of Forest Science, 48 (5): 185-201.
Kirwan, M.C., 2011. Ecological embeddedness: An interrogation and refinement of the concept within the context of alternative food networks in the UK. Journal of Rural Studies, 27: 322-330. DOI: https://doi.org/10.1016/j.jrurstud.2011.03.004.
Kowalczyk, R., Kamiński, T., Borowik, T., 2021. Do large herbivores maintain open habitats in temperate forests? Forest Ecology and Management, 494: 119310. DOI: https://doi.org/10.1016/j.foreco.2021.119310.
Lisonbee, L.D., Villalba, J.J., Provenza, F.D., Hall, J.O., 2009. Tannins and self-medication: Implications for sustainable parasite control in herbivores. Behavioural Processes, 82: 184-189. DOI: https://doi.org/10.1016/j.beproc.2009.06.009.
Liu, J., Dietz, T., Carpenter, S.R., Alberti, M., Folke, C., Moran, E., Pell, A.N., Deadman, P., Kratz, T., Lubchenco, J., …; Taylor, W.W., 2007. Complexity of coupled human and natural systems. Science, 317: 1513-1516. DOI: https://doi.org/10.1126/science.114400.
Lomba, A., Moreira, F., Klimek, S., Jongman, R.H.G., Sullivan, C., Moran, J., Poux X., Honrado, J.P., Pinto-Correia, T., Plieninger, T., McCracken, D.I., 2019. Back to the future: rethinking socioecological systems underlying high nature value farmlands. Frontiers in Ecology and the Environment, 18: 36-42. DOI: https://doi.org/10.1002/fee.2116.
MacDonald, D., Crabtree, J.R., Wiesinger, G., Dax, T., Stamou, N., Fleury, P., Gutierrez Lazpita, J., Gibon, A., 2000. Agricultural abandonment in mountain areas of Europe: Environmental consequences and policy response. Journal of Environmental Management, 59: 47-69. DOI: https://doi.org/10.1006/jema.1999.0335.
Marascuilo, L.A., 1966. Large-sample multiple comparisons. Psychological Bulletin, 65 (5): 280-290. DOI: https://doi.org/10.1037/h0023189.
Marie, P.J., Ammann, P., Boivin, G., Rey, C., 2001. Mechanisms of action and therapeutic potential of strontium in bone. Calcified Tissue International, 69: 121-129. DOI: https://doi.org/10.1037/h0023189.
Mateos, G., Fondevila, G., Cámara, L., 2019. Chapter 3: The importance of the fibre fraction of the feed in non-ruminant diets. In: G. González-Ortiz, M.R. Bedford, K.E. Bach Knudsen, C.M. Courtin and H.L. Classen, ed. The value of fibre: Engaging the second brain for animal nutrition. Wageningen Academic Publishers, pp. 61-83. DOI: https://doi.org/10.3920/978-90-8686-893-3_3.
Matuszkiewicz, W., 2008a. Przewodnik do oznaczania zbiorowisk roślinnych Polski. Warszawa: Wydawnictwo Naukowe PWN, 536 pp.
Matuszkiewicz, J., 2008b. Potential natural vegetation of Poland. Warsaw: Institute of Geography and Spatial Organization, Polish Academy of Sciences (IGSO PAS). Available from: https://www.igipz.pan.pl/potential-vegetation-dge.html.
Mirek, Z., Piękoś-Mirkowa, H., Zając, A., Zając, M., 2002. Flowering plants and pteridophytes of Poland a checklist. In: Biodiversity of Poland. Kraków: W. Szafer Institute of Botany, Polish Academy of Sciences, pp. 21-188.
Moore, K.M., Barry, T.N., Cameron, P.N., Lopez-Villalobos, N., Cameron, D.J., 2003. Willow (Salix sp.) as a supplement for grazing cattle under drought conditions. Animal Feed Science and Technology, 104: 1-11. DOI: https://doi.org/10.1016/S0377-8401(02)00326-7.
Navarro, L.M., Pereira, H.M., 2012. Rewilding abandoned landscapes in Europe. Ecosystems, 15: 900-912. DOI: https://doi.org/10.1007/s10021-012-9558-7.
Nielsen, S.P., 2004. The biological role of strontium. Bone, 35: 583-588. DOI: https://doi.org/10.1016/j.bone.2004.04.026.
Orians, G.H., Heerwagen, J.H., 1992. Evolved responses to landscapes. In: J.H. Barkow, ed. The Adapted mind: Evolutionary psychology and the generation of culture. New York: Oxford University Press, pp. 555-579.
Öllerer, K., 2014. The ground vegetation management of wood-pastures in Romania – Insights in the past for conservation management in the future. Applied Ecology and Environmental Research, 12: 549-562. DOI: https://doi.org/10.15666/aeer/1202_549562.
Peterken, G.F., Game, M., 1984. Historical factors affecting the number and distribution of vascular plant species in the woodlands of Central Lincolnshire. Journal of Ecology, 72: 155-182. DOI: https://doi.org/10.2307/2260011.
Provenza, F.D., Villalba, J.J., Dziba, L.E., Atwood, S.B., Banner, R.E., 2003. Linking herbivore experience, varied diets, and plant biochemical diversity. Small Ruminant Research, 49: 257-274. DOI: https://doi.org/10.1016/S0921-4488(03)00143-3.
Rackham, O., 2006. Woodlands. New Naturalist series. London: HarperCollins, 623 pp.
Rotherham, I.D., 2013. Cultural landscapes and problems associated with the loss of tradition and custom: an introduction and overview. In: I. Rotherham, ed. Cultural severance and the environment. Environmental history, Vol. 2. Dordrecht: Springer, pp. 3-9. DOI: https://doi.org/10.1007/978-94-007-6159-9_1.
Sebek, P., Vodka, S., Bogusch, P., Pech P., Tropek, R., Weiss, M., Zimova, K., Cizek, L., 2016. Open-grown trees as key habitats for arthropods in temperate woodlands: The diversity, composition, and conservation value of associated communities. Forest Ecology and Management, 380: 172-181. DOI: https://doi.org/10.1016/j.foreco.2016.08.052.
Slotte, H., 2001. Harvesting of leaf-hay shaped the Swedish landscape. Landscape Ecology, 16: 691-702. DOI: https://doi.org/10.1023/A:1014486331464.
Sokołowski, A.W., 1993. Fitosocjologiczna charakterystyka zbiorowisk leśnych Białowieskiego Parku Narodowego. (Phytosociological characteristics of the forest communities of the Białowieża National Park). Parki Narodowe i Rezerwaty Przyrody, 12 (3): 5-188.
Sommer, R., 1997. Further cross national studies of tree form preferences. Ecological Psychology, 9: 153-160. DOI: https://doi.org/10.1207/s15326969eco0902_3.
Spracklen, B.D., Spracklen, D.V., 2020. Old-growth forest disturbance in the Ukrainian Carpathians. Forests, 11 (2): 151. DOI: https://doi.org/10.3390/f11020151.
Tefera, S., Mlambo, V., 2017. Macro and trace elements of 26 browse species growing in semi-arid grazing lands of Eastern Cape, South Africa. African Journal of Range and Forage Science, 34: 209-217. DOI: https://doi.org/10.2989/10220119.2017.1412354.
Treydte, A.C., Heitkönig, I.M.A., Prins, H.H.T., Ludwig, F., 2007. Trees improve grass quality for herbivores in African savannas. Perspectives in Plant Ecology, Evolution and Systematics, 8: 197-205. DOI: https://doi.org/10.1016/j.ppees.2007.03.001.
Tüxen, R., 1956. Die heutige potentielle natürliche vegetation als gegenstand der vegetationskartierung. Angewandte Pflanzensoziologie, 13: 5-55.
van Laer, E., Tuyttens, F.A.M., Ampe, B., Sonck, B., Moons, C.P.H., Vandaele, L., 2015. Effect of summer conditions and shade on the production and metabolism of Holstein dairy cows on pasture in temperate climate. Animal, 9: 1547-1558.
Vandermeulen, S., Ramírez-Restrepo, C.A., Beckers, Y., Claessens, H., Bindelle, J., 2018. Agroforestry for ruminants: a review of trees and shrubs as fodder in silvopastoral temperate and tropical production systems. Animal Production Science, 58 (5): 767-777. DOI: https://doi.org/10.1071/AN16434.
Wang, B., Shi, W., Miao, Z., 2015. Confidence analysis of standard deviational ellipse and its extension into higher dimensional euclidean space. PLoS One, 10: e0118537. DOI: https://doi.org/10.1371/journal.pone.0118537.
Wolański, P., Bobiec, A., Ortyl, B., Makuch-Pietraś, I., Czarnota, P., Ziobro, J., Korol, M., Havryliuk, S., Paderewski, J., Kirby, K., 2021. The importance of livestock grazing at woodland-grassland interface in the conservation of rich oakwood plant communities in temperate Europe. Biodiversity and Conservation, 30: 741-760. DOI: https://doi.org/10.1007/s10531-021-02115-9.
Zaguła, G., Zardzewiały, M., Saletnik, B., Bajcar, M., Czernicka, M., Grabek-Lejko, D., Kasprzyk, I., Puchalski, C., 2017. Effects of fertiliser use and pre-sowing seed stimulation with a magnetic field on the mineral content and yield of three varieties of sugar beet roots. Journal of Elementology, 22 (4): 1401-1414. DOI: https://doi.org/1410.5601/jelem.2017.1422.1401.1361.
Zarzycki, J., Korzeniak, J., Perzanowska, J., 2022. Impact of land use changes on the diversity and conservation status of the vegetation of mountain grasslands (Polish Carpathians). Land, 11 (2): 252. DOI: https://doi.org/10.3390/land11020252.